Fernando Gómez

Email:   fernando.gomez (at) fitoplancton.com









Gómez, F., Kiriakoulakis, K., Lara, E. 2017. Achradina pulchra, a unique dinoflagellate (Amphilothales, Dinophyceae) with a radiolarian-like endoskeleton of celestite (strontium sulfate). Acta Protozoologica, in press.


Gómez, F. 2017. A review on the synonymy of the dinoflagellate genera Oxytoxum and Corythodinium (Oxytoxaceae, Dinophyceae). Nova Hedwigia, in press.



Gómez, F. 2017. Synonymy of dinoflagellates with basket-like endoskeletons: Monaster, Amphilothus and Achradina (Amphilothales, Dinophyceae). CICIMAR Oceánides 32(2): 1-13.


Gómez, F., Qiu, D., Lopes, R.M., Lin, S. 2017. Morphological and molecular characterization of the toxic dinoflagellate Ostreopsis cf. ovata (Dinophyta) from Brazil (South Atlantic Ocean). Revista de Biología Tropical/International Journal of Tropical Biology and Conservation 65 (3): 1022-1032.



Gómez, F., Qiu, D., Lin, S. 2017. The synonymy of the toxic dinoflagellates Prorocentrum mexicanum and P. rhathymum and the description of P. steidingerae sp. nov. (Prorocentrales, Dinophyceae). Journal of Eukaryotic Microbiology 64: 668–677. (PDF) (PDF) (PDF)


Gómez, F. 2017. The function of the ocelloid and piston in the dinoflagellate Erythropsidinium  (Gymnodiniales, Dinophyceae). Journal of Phycology, 53: 629–641. (PDF) (PDF) (PDF)


Gómez, F., Wang, L., Hernández-Becerril, D.U., Lisunova, Y.O, Lopes. R.M., Lin, S. 2017. Molecular phylogeny suggests transfer of Hemidiscus into Actinocyclus cuneiformis comb. nov. (Coscinodiscales, Bacillariophyta). Diatom Research 32, 21–28. (PDF) (PDF) (PDF)


Gómez, F. 2017. Motile behaviour of the free-living planktonic ciliate Zoothamnium pelagicum (Ciliophora, Peritrichia). European Journal of Protistology 59, 65–74.




Gómez, F., Richlen, M.L., Anderson, D.M. 2017. Molecular characterization and morphology of Cochlodinium strangulatum, the type species of Cochlodinium, and Margalefidinium gen. nov. for C. polykrikoides and allied species (Gymnodiniales, Dinophyceae).  Harmful Algae 63, 32-44. (PDF) (PDF) (PDF)


Gómez, F., Wakeman, K.C., Yamaguchi, A., Nozaki, H. 2016. Molecular phylogeny of the marine planktonic dinoflagellate Oxytoxum and Corythodinium (Peridiniales, Dinophyceae). Acta Protozoologica 55: 239–248. (PDF) (PDF) (PDF) (PDF)


Gómez, F., Qiu, D., Dodge, J.D., Lopes, R.M., Lin, S. 2016. Morphological and molecular characterization of the dinoflagellate Ptychodiscus noctiluca revealed the polyphyletic nature of the order Ptychodiscales (Dinophyceae). Journal of Phycology 52, 793–805.  (PDF) (PDF)


Gómez, F., Qiu, D., Otero-Morales, E., Lopes, R.M., Lin, S. 2016. Circumtropical distribution of the epiphytic dinoflagellate Coolia malayensis (Dinophyceae): Morphology and molecular phylogeny from Puerto Rico and Brazil. Phycological Research 64, 194199. (PDF) (PDF)


Gómez, F., Simao, T.L.L., Utz, L.R.P., Lopes, R. 2016. The nature of the diatom Leptocylindrus mediterraneus (Bacillariophyceae), host of the enigmatic symbiosis with the stramenopile Solenicola setigera. Phycologia 55, 265–273. (PDF)  (PDF) (PDF)


Gómez, F., Takayama, H., Moreira, D., López-García, P. 2016. Unarmoured dinoflagellates with a small hyposome: Torodinium and Lebouridinium gen. nov. for Katodinium glaucum (Gymnodiniales, Dinophyceae). European Journal of Phycology 52, 226-241. (PDF) (PDF) (PDF)


Gómez, F., López-García, P., Takayama, H., Moreira, D. 2015. Balechina and the new genus Cucumeridinium gen. nov. (Dinophyceae), unarmoured dinoflagellates with cell covering. Journal of Phycology 51, 1088-1105.  (PDF) (PDF) (PDF)


Gómez F., Skovgaard, A. 2015. A parasite of marine rotifers: A new lineage of dinokaryotic dinoflagellates (Dinophyceae). Journal of Marine Biology, Article ID 614609, http://dx.doi.org/10.1155/2015/614609   (PDF) (PDF)


Gómez, F., Lopes, R.M. 2015. New records of the distinctive benthic dinoflagellate genus Cabra (Dinophyceae). CICIMAR Oceánides 30: 63-70.  (PDF)  (PDF) (PDF)


Gómez, F., Qiu, D., Lopes, R.M., Lin, S. 2015. Fukuyoa paulensis gen. et sp. nov., a new genus for the globular species of the ciguatera-causing dinoflagellate Gambierdiscus (Dinophyceae). PLoS ONE 10(4): e0119676. doi:10.1371/journal.pone.0119676  (PDF)


Gómez, F., Onuma, R., Artigas, L.F. Horiguchi, T. 2015. A new definition of Adenoides eludens, an unusual marine sand-dwelling dinoflagellate without cingulum, and Pseudadenoides kofoidii, gen. et comb. nov. for the species formerly known as Adenoides eludens. European Journal of Phycology  50, 125-138. (PDF) (PDF) (PDF)


Gómez F., Skovgaard, A. 2015. Molecular phylogeny of the parasitic dinoflagellate Chytriodinium within the Gymnodinium clade (Gymnodiniales, Dinophyceae). Journal of Eukaryotic Microbiology 62, 422–425.  (PDF)  (PDF) (PDF)


Gómez F., Skovgaard, A. 2015. Molecular phylogeny of Oodinium Chatton, 1912 (Dinoflagellata: Oodiniaceae), highly divergent parasitic dinoflagellates with non-dinokaryotic characters. Systematic Parasitology 90: 125-135.  (PDF) (PDF) (PDF)


Gómez, F., Horiguchi, T. 2014. Ultrastructural features of the basal dinoflagellate Ellobiopsis chattonii (Ellobiopsidae, Alveolata), parasite of copepods. CICIMAR Océanides 29 (2): 1-10. (PDF) (PDF) (PDF)


Gómez, F., Artigas, L.F. 2014. High diversity of dinoflagellates in the intertidal sandy sediments of Wimereux (NE English Channel, France). Journal of the Marine Biological Association of the United Kingdom 94, 443-457 (PDF) (PDF) (PDF)


Gómez, F. 2014. Problematic biases in the availability of molecular markers in protists: The example of the dinoflagellates. Acta Protozoologica 53, 63-75 (PDF) (PDF)


Gómez, F., Artigas, L.F. 2013. The formation of the twin resting cysts in the dinoflagellate Dissodinium pseudolunula, a parasite of copepod eggs. Journal of Plankton Research 35, 1167-1171. (PDF) (PDF)


Gómez, F. 2013. Reinstatement of the dinoflagellate genus Tripos to replace Neoceratium, marine species of Ceratium (Dinophyceae, Alveolata). CICIMAR Océanides 28, 1-22. (PDF) (PDF)


Gómez, F. 2013. Morphology and distribution of Pronoctiluca (Dinoflagellata, incertae sedis) in the Pacific Ocean. Acta Oceanologica Sinica 32, 71-76. (PDF)


Bachy, C., Gómez, F., López-García, P., Dolan, J.R., Moreira, D. 2012. Molecular phylogeny of tintinnid ciliates (Tintinnida, Ciliophora). Protist 163, 873-887. (PDF)


Gómez, F. 2012. A quantitative review of the lifestyle, habitat and trophic diversity of dinoflagellates (Dinoflagellata, Alveolata). Systematics and Biodiversity 10, 267-275. (PDF) (PDF)


Gómez, F. 2012. A checklist and classification of living dinoflagellates (Dinoflagellata, Alveolata). CICIMAR Océanides 27, 65-140. (PDF) (PDF)


Gómez, F., López-García, P., Dolan, J.R., Moreira, D. 2012. Molecular phylogeny of the marine dinoflagellate genus Heterodinium (Dinophyceae). European Journal of Phycology 47, 95-104. (PDF)


Gómez, F., Moreira, D., López-García, P. 2012. Sinophysis and Pseudophalacroma are distantly related to typical dinophysoid dinoflagellates (Dinophysales, Dinophyceae). Journal of Eukaryotic Microbiology 59, 188-190. (PDF)


Gómez, F., Moreira, D., López-García, P. 2011. Avances en el estudio de los dinoflagelados (Dinophyceae) con la filogenia molecular. Hidrobiológica 21, 343-364. (PDF) (PDF)


Gómez, F., López-García, P., Moreira, D. 2011. Molecular phylogeny of the sand-dwelling dinoflagellates Amphidiniopsis hirsuta and A. swedmarkii (Peridiniales, Dinophyceae). Acta Protozoologica 50, 255-262. (PDF) (PDF)


Gómez, F. 2011. Diversity and distribution of the dinoflagellates Brachidinium, Asterodinium and Microceratium (Brachidiniales, Dinophyceae) in the open Mediterranean Sea. Acta Botanica Croatica 70, 209-214 (PDF)


Gómez, F. 2011. First records of the dinoflagellate genus Dicroerisma (Actiniscales, Dinophyceae) in the Mediterranean Sea. Acta Adriatica 52, 149-154 (PDF)


Gómez, F., López-García, P., Moreira, D. 2011. Molecular phylogeny of dinophysoid dinoflagellates: The systematic position of Oxyphysis oxytoxoides and the Dinophysis hastata group (Dinophysales, Dinophyceae). Journal of Phycology 47, 393-406 (PDF)


Gómez, F., 2010. Diversity and distribution of noctilucoid dinoflagellates (Noctilucales, Dinophyceae) in the open Mediterranean Sea. Acta Protozoologica 49, 365-372 (PDF) (PDF)


Gómez, F., Moreira, D., Benzerara, K., López-García, P. 2010. Solenicola setigera is the first characterized member of the abundant and cosmopolitan uncultured marine stramenopile group MAST-3. Environmental Microbiology 13, 293-302 (PDF)


Gómez, F., 2010. Changes in the Mediterranean phytoplankton community related to climate warming. In Briand, F. (Ed.) Phytoplankton Response to Mediterranean Environmental Change, CIESM Workshop Monograph 40, 37-42 (PDF)


Gómez, F. 2010. A genus name for the marine species of Ceratium. Reply to Commentary by A. Calado and J.M. Huisman on Gómez, F., Moreira, D., and López-García, P. (2010). Neoceratium gen. nov., a New Genus for All Marine Species Currently Assigned to Ceratium (Dinophyceae). Protist 161, 520-522. (PDF)


Gómez, F., Souissi, S. 2010. The diatoms Odontella sinensis, Coscinodiscus wailesii and Thalassiosira punctigera in the European Atlantic: recent introductions or overlooked in the past? Fresenius Environmental Bulletin 19, 1424-1433. (PDF)


Gómez, F., Moreira, D., López-García, P. 2010. Molecular phylogeny of noctilucoid dinoflagellates (Noctilucales, Dinophyta). Protist 161, 466-478 (PDF)


Gómez, F., Moreira, D., López-García, P. 2010. Molecular phylogeny of the dinoflagellates Podolampas and Blepharocysta (Peridiniales, Dinophyceae). Phycologia 49, 212-220 (PDF)

Gómez, F., Moreira, D., López-García, P. 2010. Neoceratium gen. nov., a new genus for all marine species currently assigned to Ceratium (Dinophyceae). Protist 161, 35-54 (PDF)

Gómez, F., Moreira, D., López-García, P. 2009. Life cycle and molecular phylogeny of the dinoflagellates Chytriodinium and Dissodinium, ectoparasites of copepod eggs. European Journal of Protistology 45, 260-270 (PDF)


Gómez, F. 2009. Torodinium and Pavillardia (Gymnodiniales, Dinophyceae): two unarmoured dinoflagellates with a body extension, collected from the open Pacific Ocean. Protistology 6, 131-135 (PDF)


Gómez, F., López-García, P., Moreira, D. 2009. Molecular phylogeny of the ocelloid-bearing dinoflagellates Erythropsidinium and Warnowia (Warnowiaceae, Dinophyceae). Journal of Eukaryotic Microbiology 56, 440-445 (PDF)


Gómez, F., López-García, P., Nowaczyk, A., Moreira, D. 2009. The crustacean parasites Ellobiopsis Caullery, 1910 and Thalassomyces Niezabitowski, 1913 form a monophyletic divergent clade within the Alveolata. Systematic Parasitology 74, 65-74 (PDF)


Gómez, F., Souissi, S. 2008. The impact of the 2003 summer heat wave and the 2005 late cold wave on the phytoplankton in the north-eastern English Channel. Comptes Rendus Biologies 331, 678-685 (PDF)

Gómez, F. 2008. Erythropsidinium (Gymnodiniales, Dinophyceae) in the Pacific Ocean, a unique dinoflagellate with an ocelloid and a piston. European Journal of Protistology 44, 291-298 (PDF)

Gómez, F., 2008. Phytoplankton invasions: Comments on the validity of categorizing the non-indigenous dinoflagellates and diatoms in European Seas. Marine Pollution Bulletin 56, 620-628 (PDF)

 Gómez, F., Claustre, H., Souissi, S. 2008. Rarely reported dinoflagellates of the genera Ceratium, Gloeodinium, Histioneis, Oxytoxum and Prorocentrum (Dinophyceae) from the open southeast Pacific Ocean. Revista de Biología Marina y Oceanografía 43, 25-40 (PDF) (PDF)

Gómez, F. 2008 (2006). Historia de las investigaciones sobre dinoflagelados marinos en España. Lull 29, 307-329 (PDF)

Gómez, F., 2008. New records of the dinoflagellate genus Dicroerisma (Actiniscales, Dinophyceae) in the Pacific Ocean. Chinese Journal of Oceanology and Limnology 26, 116-120 (PDF)

Gómez, F., 2007. Gymnodinioid dinoflagellates (Gymnodiniales, Dinophyceae) in the open Pacific Ocean. Algae 22, 273-286. (PDF)

Gómez, F., Souissi, S., 2007. Unusual diatoms linked to climatic events in the northeastern English Channel. Journal of Sea Research 58, 283-290. (PDF)

Gómez, F. 2007. Trends on the distribution of ciliates in the open Pacific Ocean. Acta Oecologica 32, 188-202. (PDF) (PDF)

Gómez, F. 2007. On the consortium of the tintinnid Eutintinnus and the diatom Chaetoceros in the Pacific Ocean. Marine Biology 151, 1899-1906. (PDF)

Gómez, F., Furuya, K. 2007. Kofoidinium, Spatulodinium and other kofoidiniaceans (Noctilucales, Dinophyceae) in the Pacific Ocean. European Journal of Protistology 43, 115-124. (PDF)

Gómez, F., Claustre H., Raimbault, P., Souissi, S. 2007. Two High-Nutrient Low-Chlorophyll phytoplankton assemblages: the tropical central Pacific and the offshore Peru-Chile Current. Biogeosciences 4, 1101-1113 (PDF)

Gómez, F., 2007. Nuevos registros del dinoflagelado Craspedotella Kofoid, 1905 (Dinophyceae: Noctilucales) en el Océano Pacífico. Revista de Biología Marina y Oceanografía 42, 83-87 (PDF)

Gómez, F., 2007. Synonymy and biogeography of the dinoflagellate genus Histioneis (Dinophysiales: Dinophyceae). Revista de Biologia Tropical 55, 459-477 (PDF)

Gómez, F., 2007. Observations on an H-shaped dinoflagellate. An example of the projection of body extensions in gymnodiniacean cells. Acta Botanica Croatica 66, 75-79 (PDF) (PDF)

Gómez, F., 2007. Gynogonadinium aequatoriale gen. et sp. nov., a new dinoflagellate from the open western equatorial Pacific. Algae 22, 11-15 (PDF)

Gómez, F., Souissi, S. 2007. The distribution and life cycle of the dinoflagellate Spatulodinium pseudonoctiluca (Dinophyceae, Noctilucales) in the northeastern English Channel. Comptes Rendus Biologies 330, 231-236 (PDF)

Gómez, F. 2007. The consortium of the protozoan Solenicola setigera and the diatom Leptocylindrus mediterraneus in the Pacific Ocean. Acta Protozoologica 46, 15-24 (PDF) (PDF)


Gómez, F. 2006. The dinoflagellate genera Brachidinium, Asterodinium, Microceratium and Karenia in the open SE Pacific Ocean. Algae 21, 445-452 (PDF)

Gómez, F. 2006. Endemic and Indo-Pacific plankton in the Mediterranean Sea: A study based on dinoflagellate records. Journal of Biogeography 33, 261-270 (PDF)

Gómez, F. 2005. Histioneis (Dinophysiales, Dinophyceae) from the western Pacific Ocean. Botanica Marina 48, 421-425. (PDF)

Gómez, F., Nagahama, Y., Takayama, H., Furuya, K. 2005. Is Karenia a synonym of Asterodinium-Brachidinium? (Gymnodiniales, Dinophyceae). Acta Botanica Croatica 64, 263-274. (PDF) (PDF)

Gómez, F., Furuya, K. 2005. Leptodiscaceans (Noctilucales, Dinophyceae) from the Pacific Ocean: First records of Petalodinium and Leptodiscus beyond the Mediterranean Sea. European Journal of Protistology 41, 231-239. (PDF)

Gómez, F., Yoshimatsu, S., Furuya, K. 2005. Morphology of Brachidinium capitatum F.J.R. Taylor (Brachidiniales, Dinophyceae) collected from the western Pacific Ocean. Cryptogamie Algologie 26, 165-175. (PDF)

Gómez, F. 2005. A list of dinoflagellates in the world’s oceans. Acta Botanica Croatica 64, 129-212. (PDF) (PDF)

Gómez, F., Furuya, K., Takeda, S. 2005. Distribution of the cyanobacterium Richelia intracellularis as an epiphyte of the diatom Chaetoceros compressus in the western Pacific Ocean. Journal of Plankton Research 27, 323-330. (PDF)

Gómez, F., Nagahama, Y., Fukuyo, Y., Furuya, K. 2004. Observations on Ceratoperidinium (Dinophyceae). Phycologia 43, 416-421. (PDF)

Gómez, F., Boicenco, L. 2004. An annotated checklist of dinoflagellates in the Black Sea. Hydrobiologia 517, 43-59. (PDF)

Gómez, F., Furuya, K. 2004. New records of Scaphodinium mirabile (Dinophyceae), an unnoticed dinoflagellate in the Pacific Ocean. Phycological Research 52, 13-16. (PDF)

Gómez, F., G. Gorsky, E. Garcia-Gorriz, M. Picheral. 2004. Control of the phytoplankton distribution in the Strait of Gibraltar by wind and fortnightly tides. Estuarine Coastal and Shelf Science 59, 485-497. (PDF)

Gómez, F. 2003. New records of Asterodinium Sournia (Brachidiniales, Dinophyceae). Nova Hedwigia 77, 331-340. (PDF) (PDF)

Gómez, F. 2003. The toxic dinoflagellate Gymnodinium catenatum: an invader in the Mediterranean Sea. Acta Botanica Croatica 62, 65-72. (PDF) (PDF)

Gómez, F. 2003. The role of the exchanges through the Strait of Gibraltar on the budget of elements in the Western Mediterranean Sea: consequences of human-induced modifications. Marine Pollution Bulletin 46, 685-694. (PDF)

Gómez, F. 2003. Checklist of Mediterranean free-living dinoflagellates. Botanica Marina 46, 215-242. (PDF)

Gómez, F., Abboud-Abi Saab, M., 2003. Records of Ceratoperidinium Margalef (Dinophyceae) from the Mediterranean Sea. Vie et Milieu 53, 43-46. (PDF)

Gómez, F., Gorsky, G. 2003. Microplankton annual cycles in the Bay of Villefranche, Ligurian Sea, NW Mediterranean Sea. Journal of Plankton Research 25, 323-339. (PDF)

Gómez, F., Claustre, H. 2003. The genus Asterodinium (Dinophyceae) as a possible biological indicator of warming in the Western Mediterranean Sea. Journal of the Marine Biological Association of United Kingdom 83, 173-174. (PDF)

Gómez, F., Gorsky, G., Striby, L., Vargas, J.M., Gonzalez, N., Picheral, M., Garcia-Lafuente, J., Varela, M., Goutx, M. 2001. Small-scale temporal variations in biogeochemical features in the Strait of Gibraltar, Mediterranean side -the role of NACW and the interface oscillation. Journal of Marine Systems 30, 207-220. (PDF) (PDF)

Gómez, F., Claustre, C. 2001. Spreading of the toxic dinoflagellate Gymnodinium catenatum in the western Mediterranean Sea. Harmful Algae News 22, 1-3. (PDF

Gómez, F., Gonzalez, N., Echevarria, F., Garcia, C.M. 2000. Distribution and fluxes of dissolved nutrients in the Strait of Gibraltar and its relationships to microphytoplankton biomass. Estuarine Coastal and Shelf Science 51, 439-449. (PDF) (PDF)

Gómez, F., Echevarria, F., Garcia, C.M., Prieto, L., Ruiz, J., Reul, A., Jimenez-Gómez, F., Varela. M. 2000. Microplankton distribution in the Strait of Gibraltar: coupling between organisms and hydrodynamic structures. Journal of Plankton Research 22, 603-617. (PDF) (PDF)